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Summary
Everyone in Naruto reincarnates. If you follow the cycles back far enough, you'll eventually find the incarnations that lived at the time of the first war against Kaguya.
Kakashi and Obito's incarnations of the time, Hatake no Okami and the forest god Kamui, were two of the leading generals in that Thousand Year War. But that was a long time ago, and reincarnation has stolen much of their past lives' memories.
Not everything, though. Some things still linger.