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This fic is sort of half fusion and half crossover. I made alterations to both worlds in order to melt their eccentricities together as seamlessly as I could manage. So buckle up and I'll do my best to get you oriented.
Most of the characters you will come across are from Supernatural. Dean's backstory has been altered a bit, but I have not changed his basic character. He's still the same rough and tumble man-whore we know and love. All canon up to Supernatural S9E12 Sharp Teeth and Dark Angel S1E1 Pilot are relevant. Any changes I have made will be revealed through the course of the fic. I won't leave you hanging or wondering unless I have to for the sake of plot. I have changed as little as canon possible before those two episodes and no changes have been made to Sam or his backstory. He still grew up rough on the road, left for Stanford, was dragged back into the family business by an overprotective big brother, and watched the love of his life burn on the ceiling. Likewise with Bobby and almost every other minor character in the series. I have made a few small changes to John due to his close association with Dean as he was growing up, but I believe that nothing I have changed will result in an alteration of his basic personality, motivations, or the major events in his life.
The biggest Dark Angel contribution to this fic is the X Series transgenics, trained from birth as soldiers. Created at Manticore by mixing human DNA with various strands of animal genetic code, they are faster, stronger, tougher, and smarter than your average homo sapien. There were quite a few mistakes in the initial attempts, think animal people, and the more unusually appearing experimental models in the later batches were done purposefully for various advantageous reasons (i.e. lizard skin doesn't lose water or burn in the desert sand as easily). The show does a good job of describing these people, so I won't go any further.
Max and her X-5 unit escaped from Manticore on schedule in 2009 just before the Pulse hit and are ignorant of the supernatural world. The Pulse was a product of Lucifer's rage, released when he crawled out of the Cage and did not find his perfect vessel there waiting for him. It devastated the American economy.
As for the Angels and the Demons, Sam completed the trials, but he barely survived. All of the demons and other denizens of Hell were sucked back into the underworld and locked in. Nothing has been heard on that front since. With the crossroads demons unavailable, Dean still made a deal with Gadreel, who he thought was Ezekiel, to save his brother's life. The spell that Metatron completed with Cas' grace locked all the Angels out of Heaven, even Metatron. In a few short years their remaining grace had been used up. Many chose death over suffering life as a human, powerless and vulnerable to pain. Gadreel quickly lost the ability to suppress Sam's consciousness as his grace dwindled, used up in the process of healing the younger Winchester, and on 28 January 2014 the angel was ejected by Sam himself without assistance. Later Sam says a few choice words that feel justified at the time, but sound harsher than he intended. Dean takes off, but does not take on the Mark of Cain. I take a hard right from Supernatural canon at this point, deviating from the path established by the show's writers. Metatron didn’t devise a spell to get back upstairs before Sam and Cas hunted him down and killed him for his treachery. Cas has not been heard from since.
I am aware that there are Dark Angel books currently in print that extend canon beyond the show. However, I have not read them yet. My canon knowledge for Dark Angel is limited to the two TV seasons that aired, and the Wikia page for those episodes. Likewise, for the Supernatural novels. Anything else about both worlds that I mention is pure extrapolation or imagination on my part. If I got anything wrong, well, smushing the canon material from two shows together necessitates a few alterations. I tried to mention everything that I changed, but stuff happens.
If you need a refresher course on Supernatural canon, the wikia is very informative and can be found HERE. I did a lot of my fact checking there.
The Dark Angel wikia can be helpful as well and if you need a few reminders, it can be found HERE.
